<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE root>
<article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xmlns:ali="http://www.niso.org/schemas/ali/1.0/" article-type="research-article" dtd-version="1.2" xml:lang="en"><front><journal-meta><journal-id journal-id-type="publisher-id">Paleontological Journal</journal-id><journal-title-group><journal-title xml:lang="en">Paleontological Journal</journal-title><trans-title-group xml:lang="ru"><trans-title>Палеонтологический журнал</trans-title></trans-title-group></journal-title-group><issn publication-format="print">0031-031X</issn><issn publication-format="electronic">3034-5871</issn><publisher><publisher-name xml:lang="en">The Russian Academy of Sciences</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="publisher-id">677970</article-id><article-id pub-id-type="doi">10.31857/S0031031X24060104</article-id><article-id pub-id-type="edn">QIEOOK</article-id><article-categories><subj-group subj-group-type="toc-heading" xml:lang="en"><subject>Articles</subject></subj-group><subj-group subj-group-type="toc-heading" xml:lang="ru"><subject>Статьи</subject></subj-group><subj-group subj-group-type="article-type"><subject>Research Article</subject></subj-group></article-categories><title-group><article-title xml:lang="en">The fallow deer <italic>Dama pontica</italic> sp. nov. (Artiodactyla, Cervidae) from the Lower pleistocene of Taurida cave in the Crimea and the early history of the genus <italic>Dama</italic></article-title><trans-title-group xml:lang="ru"><trans-title>Лань Dama pontica sp. nov. (Artiodactyla, Cervidae) из нижнего плейстоцена пещеры Таврида в Крыму и ранняя история рода Dama</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author"><name-alternatives><name xml:lang="en"><surname>Vislobokova</surname><given-names>I. А.</given-names></name><name xml:lang="ru"><surname>Вислобокова</surname><given-names>И. А.</given-names></name></name-alternatives><address><country country="RU">Russian Federation</country></address><email>ivisl@paleo.ru</email><xref ref-type="aff" rid="aff1"/></contrib></contrib-group><aff-alternatives id="aff1"><aff><institution xml:lang="en">Borissiak Paleontological Institute, Russian Academy of Sciences</institution></aff><aff><institution xml:lang="ru">Палеонтологический институт им. А.А. Борисяка РАН</institution></aff></aff-alternatives><pub-date date-type="pub" iso-8601-date="2024-12-19" publication-format="electronic"><day>19</day><month>12</month><year>2024</year></pub-date><issue>6</issue><fpage>112</fpage><lpage>124</lpage><history><date date-type="received" iso-8601-date="2025-04-01"><day>01</day><month>04</month><year>2025</year></date></history><permissions><copyright-statement xml:lang="en">Copyright ©; 2024, Russian Academy of Sciences</copyright-statement><copyright-statement xml:lang="ru">Copyright ©; 2024, Российская академия наук</copyright-statement><copyright-year>2024</copyright-year><copyright-holder xml:lang="en">Russian Academy of Sciences</copyright-holder><copyright-holder xml:lang="ru">Российская академия наук</copyright-holder></permissions><self-uri xlink:href="https://journals.eco-vector.com/0031-031X/article/view/677970">https://journals.eco-vector.com/0031-031X/article/view/677970</self-uri><abstract xml:lang="en"><p>A new fallow deer species <italic>Dama pontica</italic> is described from the late Early Pleistocene of Taurida cave in the Crimea. The antlers of this medium-sized deer had a small first process at the burr and a primitively narrow proximal palmation. The morphology of <italic>D. pontica</italic> sp. nov. allows it to be referred to a phylogenetic branch of the modern Iranian fallow deer, <italic>D. mesopotamica</italic> (Brooke, 1875). The fossil remains of <italic>Dama</italic> from the Taurida cave are the oldest in Europe. The age of the Taurida fauna, according to biochronology, is estimated at 1.8–1.5 Ma. The find indicates that the divergence of the European and Mesopotamian branches of fallow deer occurred no later than 1.5 Ma. Hypotheses about the origin of <italic>Dama </italic>(s.s.) from ‘<italic>Pseudodama</italic>’ at the Early – Middle Pleistocene transition require revision.</p></abstract><trans-abstract xml:lang="ru"><p>Новый вид лани Dama pontica sp. nov. описан по материалам из нижнего плейстоцена пещеры Таврида в Крыму. Рога этого оленя средних размеров имели маленький первый отросток у розетки и примитивно узкое проксимальное веерообразное расширение. Морфология D. pontica sp. nov. позволяет отнести этот вид к филогенетической ветви современной иранской лани, D. mesopotamica (Brooke, 1875). Остатки Dama в Тавриде – древнейшие в Европе. Возраст фауны Тавриды, по данным биохронологии, оценивается в пределах 1.8–1.5 млн л. н. Находка свидетельствует, что дивергенция ветвей европейской и месопотамской ланей произошла не позже 1.5 млн л. н. Гипотезы о происхождении ланей oт ‘Pseudodama’ на рубеже раннего и среднего плейстоцена требуют пересмотра.</p></trans-abstract><kwd-group xml:lang="en"><kwd>Dama pontica sp. nov.</kwd><kwd>Cervidae</kwd><kwd>Early Pleistocene</kwd><kwd>Crimea</kwd><kwd>Taurida Cave</kwd></kwd-group><kwd-group xml:lang="ru"><kwd>Dama pontica</kwd><kwd>Cervidae</kwd><kwd>ранний плейстоцен</kwd><kwd>Крым</kwd><kwd>пещера Таврида</kwd></kwd-group><funding-group><award-group><funding-source><institution-wrap><institution xml:lang="ru">Российский научный фонд</institution></institution-wrap><institution-wrap><institution xml:lang="en">Russian Science Foundation</institution></institution-wrap></funding-source><award-id>22-14-00214</award-id></award-group></funding-group></article-meta></front><body></body><back><ref-list><ref id="B1"><label>1.</label><mixed-citation>Верещагин Н.К. Млекопитающие Кавказа. История формирования фауны. М.–Л.: Изд-во АН СССР, 1959. 704 с.</mixed-citation></ref><ref id="B2"><label>2.</label><mixed-citation>Вислобокова И.А. Ископаемые олени Евразии. М.: Наука, 1990. 208 с.</mixed-citation></ref><ref id="B3"><label>3.</label><mixed-citation>Вислобокова И.А. История большерогих оленей (Megacerini, Cervidae, Artiodactyla). М.: ГЕОС, 2012. 103 с.</mixed-citation></ref><ref id="B4"><label>4.</label><mixed-citation>Флеров К.К. Кабарги и олени. М.; Л.: Изд-во АН СССР, 1952. 225 с.</mixed-citation></ref><ref id="B5"><label>5.</label><mixed-citation>Athanassiou A. Pleistocene vertebrates from the Kyparíssia lignite mine, Megalopolis Basin, S. Greece: Rodentia, Carnivora, Proboscidea, Perissodactyla, Ruminantia // Quatern. Intern. 2018. V. 497. P. 198–221.</mixed-citation></ref><ref id="B6"><label>6.</label><mixed-citation>Athanassiou A. The fossil record of continental fossil deer (Mammalia: Artiodactyla: Cervidae) in Greece // Fossil Vertebrates of Greece. V. 2 / Ed. Vlachos E. Springer, 2022. P. 205–247.</mixed-citation></ref><ref id="B7"><label>7.</label><mixed-citation>Azzaroli A. The cervid genus Pseudodama n. g. in the Villafranchian of Tuscany // Palaeontogr. Ital. 1992. V. 79. P. 1–41.</mixed-citation></ref><ref id="B8"><label>8.</label><mixed-citation>Baker K.H., Gray H.W.I., Lister A.M. et al. Ancient and modern DNA track temporal and spatial population dynamics in the European fallow deer since the Eemian interglacial // Sci. Rep. 2024. V. 14. № 3015. P. 1–10.</mixed-citation></ref><ref id="B9"><label>9.</label><mixed-citation>Bar-Yosef O., Belmaker M. Early and Middle Pleistocene faunal and hominins dispersals through Southwestern Asia // Quatern. Sci. Rev. 2011. V. 30. P. 1318–1337.</mixed-citation></ref><ref id="B10"><label>10.</label><mixed-citation>Bartolini-Lucenti S., Cirilli O., Pandolfi L. et al. Zoogeographic significance of Dmanisi large mammal assemblage // J. Hum. Evol. 2022. V. 163. № 103125. P. 1–18.</mixed-citation></ref><ref id="B11"><label>11.</label><mixed-citation>Belmaker M. Hominin adaptability and patterns of faunal turnover in the Lower–Middle Pleistocene transition in the Levant // A Sourcebook of Paleolithic Transitions: Methods, Theories and Interpretations / Eds. Camps M., Chauhan P.R. N.Y.: Springer, 2009. P. 211–227.</mixed-citation></ref><ref id="B12"><label>12.</label><mixed-citation>Breda M. The early Middle Pleistocene fallow deer Dama roberti: new insight on species morphology from a complete postcranial skeleton from Valdemino (northwestern Italy) // Geol. J. 2015. V. 50. P. 257–270.</mixed-citation></ref><ref id="B13"><label>13.</label><mixed-citation>Breda M., Kahlke R.-D., Lister A.M. New results on cervids from the Early Pleistocene site of Untermassfeld // The Pleistocene of Untermassfeld near Meiningen (Thüringen, Germany). Part 4 / Ed. Kahlke R.-D. Monogr. Röm.-German. Zentralmuseums. 2020. Bd 40. H. 4. P. 1197–1249.</mixed-citation></ref><ref id="B14"><label>14.</label><mixed-citation>Breda M., Lister A.M. Dama roberti, a new species of deer from the early Middle Pleistocene of Europe, and the origins of modern fallow deer // Quatern. Sci. Rev. 2013. V. 69. № 1. P. 155–167.</mixed-citation></ref><ref id="B15"><label>15.</label><mixed-citation>Brooke V. On a new species of deer from Mesopotamia // Proc. Zool. Soc. London. 1875. P. 261–266.</mixed-citation></ref><ref id="B16"><label>16.</label><mixed-citation>Bukhsianidze M. Catalogue of the holotypes of fossil land mammals in the Georgian National Museum. Tbilisi: Georgian National Museum, 2015. 103 p.</mixed-citation></ref><ref id="B17"><label>17.</label><mixed-citation>Cherin M., Breda M., Esattore B. et al. A Pleistocene Fight Club revealed by the palaeobiological study of the Dama-like deer record from Pantalla (Italy) // Sci. Rep. 2022. V. 12. № 13898. Р. 1–11.</mixed-citation></ref><ref id="B18"><label>18.</label><mixed-citation>Croitor R. Early Pleistocene small-sized deer of Europe // Hellen. J. Geosci. 2006. V. 41. № 1. P. 89–117.</mixed-citation></ref><ref id="B19"><label>19.</label><mixed-citation>Croitor R. Plio-Pleistocene Deer of Western Palearctic: Taxonomy, Systematics, Phylogeny. Chişinău: Inst. Zool. Acad. Sci. Moldova, 2018. 140 p.</mixed-citation></ref><ref id="B20"><label>20.</label><mixed-citation>Deng Tao. Chinese Neogene mammal biochronology // Vertebr. PalAsiat. 2006. V. 44. № 2. P. 143–163.</mixed-citation></ref><ref id="B21"><label>21.</label><mixed-citation>Di Stefano G. The Mesopotamian fallow deer (Dama, Artiodactyla) in the Middle East Pleistocene // N. Jb. Geol. Paläontol. Abh. 1996. Bd 199. H. 3. P. 295–322.</mixed-citation></ref><ref id="B22"><label>22.</label><mixed-citation>Di Stefano G., Petronio C. Origin of and relationships among Dama-like cervids in Europe // N. Jb. Geol. Paläontol. Abh. 1998. Bd 207. H. 1. P. 37–55.</mixed-citation></ref><ref id="B23"><label>23.</label><mixed-citation>Di Stefano G., Petronio C. Systematics and evolution of the Eurasian Plio-Pleistocene tribe Cervini (Artiodactyla, Mammalia) // Geol. Romana. 2002. V. 36. P. 311–334.</mixed-citation></ref><ref id="B24"><label>24.</label><mixed-citation>Dong W. Reconsideration of the systematics of the Early Pleistocene Cervavitus (Cervidae, Artiodactyla, Mammalia) // Estud. Geol. 2011. V. 67. № 2. P 603–611.</mixed-citation></ref><ref id="B25"><label>25.</label><mixed-citation>Groves C., Grubb P. Ungulate Taxonomy. Baltimore: The John Hopkins Univ. Press, 2011. 310 p.</mixed-citation></ref><ref id="B26"><label>26.</label><mixed-citation>Haltenorth T. Beitrag zur Kenntnis des mesopotamischen Damhirsches Cervus (Dama) mesopotamicus Brook und zur Stammes- und Verbreitungs-Geschichte der Damhirsche allgemein // Säugt. Mitt. 1959. Bd 7. S. 1–89.</mixed-citation></ref><ref id="B27"><label>27.</label><mixed-citation>Hassanin A., Delsuc F., Ropiquet A. et al. Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes // C.R. Soc. Biol. 2012. V. 335. P. 32–50.</mixed-citation></ref><ref id="B28"><label>28.</label><mixed-citation>Heintz E. Les cervidés villafranchiens de France et d’Espagne // Mém. Mus. Nat. Hist. Natur. Sér. C. 1970. T. 22. P. 1–303.</mixed-citation></ref><ref id="B29"><label>29.</label><mixed-citation>Kahlke H.-D. Die Cerviden-Reste aus dem Unterpleistozän von Untermaßfeld // Das Pleistozän von Untermaßfeld bei Meiningen (Thüringen). T. 1 / Ed. Kahlke R.-D. Monogr. Röm.-German. Zentralmuseums. 1997. Bd 40. H. 1. S. 181–275.</mixed-citation></ref><ref id="B30"><label>30.</label><mixed-citation>Kahlke H.-D. Neufunde von Cerviden-Resten aus dem Unterpleistozän von Untermaßfeld // Das Pleistozän von Untermaßfeld bei Meiningen (Thüringen). T. 2 / Ed. Kahlke R.-D. Monogr. Röm.-German. Zentralmuseums. 2001. Bd 40. H. 2. S. 461–482.</mixed-citation></ref><ref id="B31"><label>31.</label><mixed-citation>Karastoyanova N., Gorczyk J., Spassov N. The natural history of the fallow deer, Dama dama (Linnaeus, 1758) in Bulgaria in prehistory and new evidence for the existence of an autochthonous Holocene population in the Balkans // Int. J. Osteoarchaeol. 2020. V. 30. P. 616–628.</mixed-citation></ref><ref id="B32"><label>32.</label><mixed-citation>Leonardi G., Petronio C. The fallow deer of European Pleistocene // Geol. Romana. 1976. V. 25. P. 1–67.</mixed-citation></ref><ref id="B33"><label>33.</label><mixed-citation>Lister A.M. The morphological distinction between bones and teeth of fallow deer (Dama dama) and red deer (Cervus elaphus) // Int. J. Osteoarchaeol. 1996. V. 6. P. 119–143.</mixed-citation></ref><ref id="B34"><label>34.</label><mixed-citation>Lister A.M. Cervidae // The Lower Paleolithic Site of Holon, Israel / Eds Chazan M., Horwitz L.K. Cambridge: Harvard Univ. Press, 2007. P. 111–121.</mixed-citation></ref><ref id="B35"><label>35.</label><mixed-citation>Lister A.M., Edwards C.J., Nock D.A.W. et al. The phylogenetic position of the ‘giant deer’ Megaloceros giganteus // Nature. 2005. V. 438. № 8. P. 850–853.</mixed-citation></ref><ref id="B36"><label>36.</label><mixed-citation>Lumley H. de, Kahlke H.-D., Moigne A.-M., Moullé P.-E. Les faunes de grands mammifères de la grotte du Vallonnet Roquebrune-Cap-Martin, Alpes-Maritimes // L’Anthropologie. 1988. V. 92. № 2. P. 465–469.</mixed-citation></ref><ref id="B37"><label>37.</label><mixed-citation>Marzban Abbasabadi B., Ghojoghi F., Rahmati S., Hajian O. Morphological variations in the skulls of male and female Persian fallow deer (Dama dama mesopotamica) // Iran. J. Vet. Med. 2020. V. 15. № 2. P. 221–233.</mixed-citation></ref><ref id="B38"><label>38.</label><mixed-citation>Masini F., Sala B. Large and small-mammal distribution patterns and chronostratigraphic boundaries from the Late Pliocene to the Middle Pleistocene of the Italian peninsula // Quatern. Intern. 2007. V. 160. P. 43–56.</mixed-citation></ref><ref id="B39"><label>39.</label><mixed-citation>Masseti M., Mertzanidou D. Dama dama. The IUCN Red List of Threatened Species. 2008. e.T42188A10656554. P. 1–10.</mixed-citation></ref><ref id="B40"><label>40.</label><mixed-citation>Masseti M., Vernesi C. Historic zoology of the European fallow deer, Dama dama dama: Evidence from biogeography, archaeology and genetics // Deer and People / Eds Baker K., Carden R., Madgwick R. Oxford: Windgather Press, 2014. P. 13–22.</mixed-citation></ref><ref id="B41"><label>41.</label><mixed-citation>Mecozzi B., Sardella R., Breda M. Late Early to late Middle Pleistocene mediumsized deer from the Italian Peninsula: implications for taxonomy and biochronology // Palaeobiodiv. Palaeoenv. 2024. V. 104. P. 191–215.</mixed-citation></ref><ref id="B42"><label>42.</label><mixed-citation>Moncel M.H., Santagata C., Pereira A. et al. The origin of early Acheulean expansion in Europe 700 ka ago: new findings at Notarchirico (Italy) // Sci. Rep. 2020. V. 10. № 1. P. 1–16.</mixed-citation></ref><ref id="B43"><label>43.</label><mixed-citation>Pfeiffer T. Die fossilen Damhirsche von Neumark-Nord (Sachsen-Anhalt) – Dama dama geiselana n. ssp. // Eiszeit. Gegenw. 1998. Bd 48. S. 72–86.</mixed-citation></ref><ref id="B44"><label>44.</label><mixed-citation>Pfeiffer T. Die Stellung von Dama (Cervidae, Mammalia) im System plesiometacarpaler Hirsche des Pleistozäns // Cour. Forsch.-Inst. Senckenb. 1999. Bd 211. S. 1–218.</mixed-citation></ref><ref id="B45"><label>45.</label><mixed-citation>Pfeiffer T. The position of Dama (Cervidae, Mammalia) in the system of fossil and living deer from Europe-Phylogenetical analysis based on the postcranial skeleton // Quatern. Hors-sér. 2005. V. 2. P. 39–57.</mixed-citation></ref><ref id="B46"><label>46.</label><mixed-citation>Pfeiffer-Deml T. Deer from the Pliocene site of Bad Deutsch-Altenburg 26 (Lower Austria, Leithagebirge): Conclusions based on skeletal morphology // Ann. Naturhist. Mus. Wien. Ser. A. 2016. V. 118. P. 133–173.</mixed-citation></ref><ref id="B47"><label>47.</label><mixed-citation>Pfeiffer-Deml T. The fossil fallow deer Dama geiselana (Cervidase, Mammalia, upgrade to species level) in the context of migration and local extinctions of fallow deer in the Late and Middle Pleistocene of Europe // Paläontol. Z. 2018. V. 92. P. 681–713.</mixed-citation></ref><ref id="B48"><label>48.</label><mixed-citation>Pitra C., Fickel J., Meijaard E., Groves C.P. Evolution and phylogeny of old world deer // Mol. Phylog. Evol. 2004. V. 33. № 3. P. 880–895.</mixed-citation></ref><ref id="B49"><label>49.</label><mixed-citation>Qiu Z. Some mammalian fossils from the Pliocene of Inner Mongolia and Gansu (Kansu) // Vertebr. PalAsiat. 1979. V. 17. № 3. P. 222–235.</mixed-citation></ref><ref id="B50"><label>50.</label><mixed-citation>Rabinovich R., Biton R. The Early-Middle Pleistocene faunal assemblages of Gesher Benot Ya‘aqov: Inter-site variability // J. Hum. Evol. 2011. V. 60. P. 357–374.</mixed-citation></ref><ref id="B51"><label>51.</label><mixed-citation>Rabinovich R., Gaudzinski-Windheuser S., Goren-Inbar N. Systematic butchering of fallow deer (Dama) at the early Middle Pleistocene Acheulian site of Gesher Benot Ya’aqov (Israel) // J. Hum. Evol. 2008. V. 54. P. 134–149.</mixed-citation></ref><ref id="B52"><label>52.</label><mixed-citation>Randi E., Mucci N., Claro-Hergeta F. et al. A mitochondrial DNA control region phylogeny of the Cervinae: speciation in Cervus and implications for conservation // Anim. Conserv. 2001. V. 4. P. 1–11.</mixed-citation></ref><ref id="B53"><label>53.</label><mixed-citation>Rivals F. Les petits bovidés (Caprini et Rupicaprini) pléistocènes dans le bassin méditerranéen et le Caucase. Étude paléontologique, biostratigraphique, archéozoologizue et paléoécologique // Brit. Archaeol. Rep. Intern. Ser. 2004. V. 1327. P. 1–252.</mixed-citation></ref><ref id="B54"><label>54.</label><mixed-citation>Sickenberg O. Eine Saugetierfauna des tieferen Bihariums aus dem Becken von Megalopolis (Peloponnes, Griehenland) // Anal. Geol. Pays Hellen. Ser. 1. 1976. V. 27. P. 25–73.</mixed-citation></ref><ref id="B55"><label>55.</label><mixed-citation>Teilhard de Chardin P., Trassaert M. The Pliocene Camelidae, Giraffidae, and Cervidae of South Eastern Shansi // Palaeontol. Sin. N. Ser. C. 1937. № 1. P. 1–56.</mixed-citation></ref><ref id="B56"><label>56.</label><mixed-citation>Tsuboi M., Kopperud B.T., Matschiner M. et al. Antler allometry, the Irish Elk and Gould revisited // Evol. Biol. 2024. V. 51. P. 149–165.</mixed-citation></ref><ref id="B57"><label>57.</label><mixed-citation>Van der Made J. Ungulates from Atapuerca TD6 // J. Hum. Evol. 1999. V. 37. P. 389–413.</mixed-citation></ref><ref id="B58"><label>58.</label><mixed-citation>Van der Made J. The latest Early Pleistocene giant deer Megaloceros novocarthaginiensis n. sp. and the fallow deer Dama cf. vallonnetensis from Cueva Victoria (Murcia, Spain) // Mastia. 2015. V. 11–13. P. 269–323.</mixed-citation></ref><ref id="B59"><label>59.</label><mixed-citation>Van der Made J., Rodriguez-Alba J.J., Martos J.A. et al. The fallow deer Dama celiae sp. nov. with two-pointed antlers from the Middle Pleistocene of Madrid, a contemporary of humans with Acheulean technology // Archaeol. Antropol. Sci. 2023. V. 15. № 41. P. 1–35.</mixed-citation></ref><ref id="B60"><label>60.</label><mixed-citation>Van der Made J., Torres T., Ortiz J.E. et al. The new material of large mammals from Azokh and comments on the older collections // Azokh Caves and the Transcaucasian Corridor / Eds Fernández-Jalvo Y. et al. Springer, 2016. P. 117–162.</mixed-citation></ref><ref id="B61"><label>61.</label><mixed-citation>Vekua A. Die Wirbeltirfauna des Villafranchium von Dmanisi und ihre biostratigraphische Bedeutung // Jb. Röm.-German. Zentralmuseums. 1995. Bd 42. S. 77–180.</mixed-citation></ref><ref id="B62"><label>62.</label><mixed-citation>Vereshchagin N.K., Baryshnikov G.F. Quaternary mammalian extinction in Northern Eurasia // Quaternary Extinctions / Eds. Martin P.S., Klein R.G. Tucson: The Univ. Arizona Press, 1984. P. 483–516.</mixed-citation></ref><ref id="B63"><label>63.</label><mixed-citation>Werner N.Y., Rabiei A., Saltz D. et al. Dama mesopotamica. The IUCN Red List of Threatened Species 2015: e.T6232A97672550. P. 1–15.</mixed-citation></ref><ref id="B64"><label>64.</label><mixed-citation>Woodburne M.O., Tedford R.H., Lindsay E.H. North China Neogene biochronology: A Chinese standard // Fossil Mammals of Asia / Eds Wang X., Flynn L.J., Fortelius M. N.Y.: Columbia Univ. Press, 2013. P. 91–123.</mixed-citation></ref></ref-list></back></article>
