<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE root>
<article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xmlns:ali="http://www.niso.org/schemas/ali/1.0/" article-type="research-article" dtd-version="1.2" xml:lang="en"><front><journal-meta><journal-id journal-id-type="publisher-id">Doklady Biological Sciences</journal-id><journal-title-group><journal-title xml:lang="en">Doklady Biological Sciences</journal-title><trans-title-group xml:lang="ru"><trans-title>Доклады Российской академии наук. Науки о жизни</trans-title></trans-title-group></journal-title-group><issn publication-format="print">2686-7389</issn><issn publication-format="electronic">3034-5057</issn><publisher><publisher-name xml:lang="en">The Russian Academy of Sciences</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="publisher-id">697706</article-id><article-id pub-id-type="doi">10.7868/S3034543X25050029</article-id><article-categories><subj-group subj-group-type="toc-heading" xml:lang="en"><subject>Articles</subject></subj-group><subj-group subj-group-type="toc-heading" xml:lang="ru"><subject>Статьи</subject></subj-group><subj-group subj-group-type="article-type"><subject>Research Article</subject></subj-group></article-categories><title-group><article-title xml:lang="en">REST AND SLEEP IN THE HIPPOPOTAMUS (<italic>HIPPOPOTAMUS AMPHIBIOUS</italic>)</article-title><trans-title-group xml:lang="ru"><trans-title>ПОКОЙ И СОН У ОБЫКНОВЕННОГО ГИППОПОТАМА (<italic>HIPPOPOTAMUS AMPHIBIUS</italic>)</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author"><name-alternatives><name xml:lang="en"><surname>Lyamin</surname><given-names>O. I</given-names></name><name xml:lang="ru"><surname>Лямин</surname><given-names>О. И</given-names></name></name-alternatives><email>oilyamin@yahoo.com</email><xref ref-type="aff" rid="aff1"/></contrib><contrib contrib-type="author"><name-alternatives><name xml:lang="en"><surname>Siegel</surname><given-names>J. M</given-names></name><name xml:lang="ru"><surname>Сигал</surname><given-names>Д. М</given-names></name></name-alternatives><xref ref-type="aff" rid="aff2"/></contrib></contrib-group><aff-alternatives id="aff1"><aff><institution xml:lang="en">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences</institution></aff><aff><institution xml:lang="ru">Федеральное государственное бюджетное учреждение науки Институт проблем экологии и эволюции им. А.Н. Северцова Российской академии наук</institution></aff></aff-alternatives><aff-alternatives id="aff2"><aff><institution xml:lang="en">University of California in Los Angeles</institution></aff><aff><institution xml:lang="ru">Калифорнийский университет в г. Лос-Анджелес</institution></aff></aff-alternatives><pub-date date-type="pub" iso-8601-date="2025-10-15" publication-format="electronic"><day>15</day><month>10</month><year>2025</year></pub-date><volume>524</volume><issue>1</issue><issue-title xml:lang="en">VOL 524, NO1 (2025)</issue-title><issue-title xml:lang="ru">ТОМ 524, №1 (2025)</issue-title><fpage>502</fpage><lpage>508</lpage><history><date date-type="received" iso-8601-date="2025-12-04"><day>04</day><month>12</month><year>2025</year></date></history><permissions><copyright-statement xml:lang="en">Copyright ©; 2025, Russian Academy of Sciences</copyright-statement><copyright-statement xml:lang="ru">Copyright ©; 2025, Российская академия наук</copyright-statement><copyright-year>2025</copyright-year><copyright-holder xml:lang="en">Russian Academy of Sciences</copyright-holder><copyright-holder xml:lang="ru">Российская академия наук</copyright-holder><ali:free_to_read xmlns:ali="http://www.niso.org/schemas/ali/1.0/" start_date="2026-10-15"/></permissions><self-uri xlink:href="https://journals.eco-vector.com/2686-7389/article/view/697706">https://journals.eco-vector.com/2686-7389/article/view/697706</self-uri><abstract xml:lang="en"><p>Rest and sleep states of an adult pair of hippopotamuses (Hippopotamus amphibius) and their two-month-old calf are described in a zoo (Tampa, USA). During the day, the animals spent an equal time on land and in water; at night, they spent on average 76% of their time in water. Rest occupied 48-53% of the 24-hour period (11.4–12.8 h). On land, the animals rested in a lying position (18% of rest in the calf); in water – lying, sitting, or standing on the bottom with nostrils, eyes, and ears above the surface (80% in the female), or submerged and lying on the bottom (29% in the calf). All displayed eye jerks and twitches resembling REM sleep. On land, 98% of breathing pauses lasted &lt;30 sec. In water, they ranged from 4 to 145 sec, with most lasting &lt;1 min. The male's eyes were closed more often (up to 72%), the female's partially open (84%). Slow wave sleep in the hippopotamus is likely bilaterally symmetrical, as in terrestrial mammals. The female's sleep was more fragmented, less deep, and vigilant due to calf care.</p></abstract><trans-abstract xml:lang="ru"><p>По видеозаписям описаны особенности покоя и сна взрослой пары гиппопотамов (Hippopotamus amphibius) и их двухмесячного детеныша в зоопарке (г. Тампа, США). Днем животные проводили равное время на суше и в воде, ночью — в среднем 76% (9.2 час) в воде. На покой приходилось 48-53% времени суток (11.4-12.8 час). На суше гиппопотамы отдыхали лежа (в среднем 18% покоя у детеныша), в воде — лежа, сидя или стоя на дне с выставленными над поверхностью ноздрами, глазами и ушами (у самки 80%) или погружаясь и ложась на дно (у детеныша 29%). В покое наблюдались вздрагивания глаз, ушей и ноздрей, напоминавшие REM сон. На суше 98% дыхательных пауз были &lt;30 с. В воде они варьировали от 4 до 145 с, большинство длилось меньше минуты. У самца глаза были чаще закрыты (до 72% времени), у самки — приоткрыты (84%). Медленноволновый сон у гиппопотамов, вероятно, билатерально симметричный, как у наземных млекопитающих. Сон у самки более фрагментированный, менее глубокий и более бдительный из-за заботы о детеныше.</p></trans-abstract><kwd-group xml:lang="en"><kwd>rest</kwd><kwd>sleep</kwd><kwd>slow wave sleep</kwd><kwd>REM sleep</kwd><kwd>vigilance</kwd><kwd>common hippopotamus</kwd><kwd><italic>Hippopotamus amphibius</italic></kwd><kwd>ungulates</kwd><kwd>semiaquatic mammals</kwd><kwd>cetaceans</kwd></kwd-group><kwd-group xml:lang="ru"><kwd>покой</kwd><kwd>сон</kwd><kwd>медленноволновый сон</kwd><kwd>REM сон</kwd><kwd>бдительность</kwd><kwd>обыкновенный гиппопотам</kwd><kwd><italic>Hippopotamus amphibius</italic></kwd><kwd>копытные</kwd><kwd>полуводные млекопитающие</kwd><kwd>китообразные</kwd></kwd-group><funding-group><funding-statement xml:lang="ru">Исследование было поддержано NIH (США)</funding-statement></funding-group></article-meta></front><body></body><back><ref-list><ref id="B1"><label>1.</label><mixed-citation>Ковальзон В. М. Нейробиология бодрствования и сна. М.: УРС. 2024.</mixed-citation></ref><ref id="B2"><label>2.</label><mixed-citation>Лямин О.Н., Косенко П.О., Высоцкий А.Л., и др. // Доклады Академии наук. 2012. Т. 444. N. 4. C. 461–465.</mixed-citation></ref><ref id="B3"><label>3.</label><mixed-citation>Лямин О.Н., Сигал Д.М., Назаренко Е.А., и др. // Доклады Российской академии наук. Науки о жизни. 2023. Т. 512. C. 480–485.</mixed-citation></ref><ref id="B4"><label>4.</label><mixed-citation>Elringham S.K. The Hippos. London: Academic Press. 1999.</mixed-citation></ref><ref id="B5"><label>5.</label><mixed-citation>Lewison R.L., Carter J. // Ecological Modelling. 2004. V. 171. N. 1–2. P. 127–138.</mixed-citation></ref><ref id="B6"><label>6.</label><mixed-citation>Mekonen S., Hallemartam B. // American J. Of Scientific And Industrial Research. 2016. V. 7. N. 2. P. 41–49.</mixed-citation></ref><ref id="B7"><label>7.</label><mixed-citation>Bempah G., Osei M., Gyimath A., et al. // bioRxiv 2021.08.30.458240; doi:https://doi.org/10.1101/2021.08.30.458240</mixed-citation></ref><ref id="B8"><label>8.</label><mixed-citation>Fernandez E.J., Ramirez M., Hawkes N.C. // Animals (Basel). 2020. V. 10. N. 6. P. 1022.</mixed-citation></ref><ref id="B9"><label>9.</label><mixed-citation>Ртузьболо Г.Р., Lyamin O.I., Siegel J.M., et al. // Behavioral Brain Research. 2009. V. 201. P. 80–87.</mixed-citation></ref><ref id="B10"><label>10.</label><mixed-citation>Ruckebusch Y. // Anim Behav. 1972. V. 20, N. 4:637–643.</mixed-citation></ref><ref id="B11"><label>11.</label><mixed-citation>Lyamin O.I., Siegel J.M., Nazarenko E.A., et al. // Sleep. 2022. V. 11. N. 45(7):zsab199.</mixed-citation></ref><ref id="B12"><label>12.</label><mixed-citation>Kendall-Bar J.M., Williams T.M., Mukherji R., et al. // Science. 2023. V. 380. N. 6642. P. 260–265.</mixed-citation></ref><ref id="B13"><label>13.</label><mixed-citation>Tobler I. // Sleep. 1992. V. 15. N. 1. P. 1–12.</mixed-citation></ref><ref id="B14"><label>14.</label><mixed-citation>Tobler I., Schwierin B. // J. Sleep Res. V1996. 5(1). P. 21–32.</mixed-citation></ref><ref id="B15"><label>15.</label><mixed-citation>Lima C. J. // Anim. Behav. 2005. V. 70. P. 723–736.</mixed-citation></ref></ref-list></back></article>
